CHAPTER 21    BLOOD VESSELS

 

Blood vessels are a series of branching tubes that carry blood from heart to tissues and back to the heart.

 

1. Arteries--carry blood AWAY from heart to tissues   

   a. Large elastic conducting arteries

   b. Medium muscular distributing arteries

   c. Small arteries

2. Arterioles

3. Capillaries--in tissues, these are where exchange takes place

4. Venules--smallest veins, formed as capillaries unite

5. Veins--carry blood from tissues back to the heart

 

STRUCTURE OF BLOOD VESSELS

 

The lumen is the hollow center of a blood vessel (or any hollow organ).

A. ARTERIES--the wall is in 3 layers:

   1. Tunica intima or interna--inner coat

      a. Lining of endothelium—lines all blood vessels and the heart. This is the ONLY

           tissue blood should come in contact with

      b. Basement membrane

      c. Internal elastic lamina--layer of elastic tissue

 

   2. Tunica media--middle coat--this is the thickest layer and consists of elastic fibers and smooth muscle

 

   3. Tunica adventitia or externa--outer coat--composed of elastic and collagen fibers

 

 

Due mostly to the structure of the tunica media, arteries have 2 essential properties:                    

   1. Elasticity--as blood is forced from the heart into large arteries, the elastic tissue in the wall first stretches without tearing and then snaps back (elastic recoil) to push the blood onward. This helps keep the blood moving while the ventricles are relaxed.

 

   2. Contractility--smooth muscle in the wall can contract to narrow the lumen (vasoconstriction). The opposite effect, vasodilation, occurs when the smooth muscle relaxes. Sympathetic nerves  control the muscle in the walls of blood vessels, but these smooth muscle fibers can also change their degree of contraction due to local factors.   

 

 

The largest arteries are elastic (conducting) arteries. Walls are relatively thin and tunica media contains large numbers of elastic fibers and relatively little smooth muscle. Their elastic fibers stretch & then recoil, pushing blood onwards while the ventricles are relaxed. Includes:

        Aorta                                Subclavian                   Pulmonary

        Brachiocephalic                Vertebral

        Common carotid               Common iliac

              

 

 Medium-sized arteries branch off the elastic conducting arteries and are muscular (distributing) arteries. Their walls are relatively thick and contain more smooth muscle and less elastic fibers. They are capable of greater vasoconstriction. Includes:

   Axillary                          Mesenteric

   Brachial                         Femoral

   Radial                            Popliteal

   Intercostal                     Tibial

   Splenic

  

 

B. ARTERIOLES are very small arteries that deliver blood to capillaries. As arterioles branch off an artery, they have smooth muscle and a few elastic fibers in the tunica media. These gradually taper away as the arteriole becomes smaller, leaving mostly the endothelium and a few smooth muscle fibers by the time the arteriole connects to the capillaries.

 

Arterioles play a key role in regulating blood flow into capillaries. Vasoconstriction of arterioles decreases blood flow into capillaries; vasodilation increases flow. A change in the diameter of a large number of arterioles at once will also affect blood pressure.

 

C. CAPILLARIES--microscopic vessels that connect arterioles and venules. They are found near almost every cell in the body, but are more plentiful in tissues with high metabolic activity (liver, kidneys, muscles, CNS). A few tissues lack capillaries (sheets of covering and lining epithelium such as epidermis and  epithelial linings of viscera, cornea, lens, cartilage).

 

Capillaries permit the exchange of nutrients and wastes between blood and tissue cells. The walls consist of a single layer of endothelial cells and a basement membrane, so substances must cross only one cell layer. Walls of most other blood vessels are too thick to allow exchange in the time available as the blood flows through them. (Remember, blood is always moving.) Some exchange does occur across the walls of the very first part of venules.

 

Most tissues contain many capillaries, often in the form of extensive branching networks, to increase the surface area for diffusion and allow the rapid exchange of materials. However, all capillaries in the network (also known as the capillary bed) may not be filled with flowing blood at all times.  When metabolic needs of the tissue are low, blood flows through only a portion of the network. More of the network fills with blood as the tissue becomes active. This is made possible by:

 

1. Metarterioles--special vessels with smooth muscle fibers in their walls. They emerge from an arteriole and enter the capillary network. The distal portion of the metarteriole, known as a thoroughfare channel, will empty into a venule. Metarterioles open when blood flow through the capillaries of the network needs to be reduced. With the metarteriole open, a relatively small amount of blood still flows through its capillaries and most of the blood passes straight through to the venule. When the tissue needs the maximum amount of blood, most of the metarterioles constrict so that more blood flows out into the entire capillary network. 

 

2. Precapillary sphincters--capillaries in networks have rings of smooth muscle at their origin. These sphincters open (relax) when the tissue is active to allow more blood to enter  and close (contract) when less blood is needed.

 

Remember, precapillary sphincters relax when metarteriole muscle contracts. Precapillary sphincters contract when metarteriole muscle relaxes. In a process called vasomotion, there is alternation between precapillary sphincter contraction and metarteriole muscle contraction in an area, so that blood flow through the capillary network is intermittent. Unless there is great demand, blood is usually flowing through only about 25% of the capillaries in an area at one time.

 

TYPES OF CAPILLARIES

 

1. Continuous capillaries—plasma membranes of endothelial cells form  tubes interrupted only by intercellular clefts—gaps between endothelial cells.

                Skeletal muscle

                Smooth muscle

                Connective tissue       

                Lungs

 

2. Fenestrated capillaries—plasma membranes have tiny openings called fenestrations or pores. The size of these pores varies.

                  Kidney

                  Small intestine

                  Choroid plexuses of brain ventricles

                  Ciliary processes of eyes

                  Endocrine glands

 

3. Sinusoids—these capillaries are wider and more twisting than regular capillaries. Their endothelial cells contain relatively large intercellular clefts and also very large fenestrations (pores).  The basement membrane may be incomplete or absent. Walls of sinusoids may contain phagocytic cells which remove bacteria, worn-out blood cells, and debris from the blood as it flows through.

        Liver                                         Spleen

        Anterior pituitary                      Parathyroids

        Red bone marrow       

 

D. VENULES are formed when capillaries unite. The walls of the smallest venules are thin with no tunica externa—this is the place where phagocytic WBC often leave the blood to migrate to trouble spots. As venules become larger, they begin to develop a thicker wall with all 3 layers. Larger venules unite to form veins.

 

 

E. VEINS are composed of the same 3 coats as arteries but characteristics vary:

 

   1. Tunica interna (intima) is thinner with no internal elastic lamina

 

   2. Tunica media is much thinner, with only very small numbers of smooth muscle

       and elastic fibers. Although the amount of smooth muscle present is small,  it

        is enough to allow veins to alter their diameter to some extent.

 

   3. Tunica externa (adventitia) is thicker. This is the thickest layer of a vein wall

        and contains more fibrous CT (collagen and elastic fibers).

 

Compared to arteries that carry similar amounts of blood, vein walls are thinner and softer and the lumen is larger in diameter.                                      

 

In certain locations of the body, veins are modified to form venous (vascular) sinuses. The walls consist of endothelium supported by surrounding dense CT. There is no tunica media or tunica externa and no smooth muscle. Venous sinuses are found in the brain and heart.

 

Blood in veins has a much lower pressure than blood in arteries. Most veins contain valves, which are flaps (cusps) from the tunica intima, to maintain flow in the proper direction. Varicose veins occur when valves leak. Blood pools in that part of the vein and stretches the wall. This is most frequently seen in leg veins, where gravity works against the flow of blood in veins, but can occur in any vein.

 

Most tissues receive blood from more than one artery. The union of the branches of 2 or more arteries supplying the same body region is an anastomosis. Anastomoses provide alternate routes for blood to reach a tissue or organ--this is called collateral circulation. Arteries that do not anastomose are called end arteries.

 

 

 

BLOOD DISTRIBUTION

 

At rest the blood is distributed as follows:

  64% in systemic veins & venules

   7% in systemic capillaries

  13% in systemic arteries & arterioles

   7% in heart

   9% in pulmonary (lung) vessels

 

Systemic veins and venules are known as blood reservoirs. A good bit of the blood contained in them can be quickly diverted to other vessels if needed. The veins constrict and the blood flows to where it is needed:

   Exercise—skeletal muscle

   Hemorrhage—to circulation

Major blood reservoirs are the veins of the liver, spleen, and skin.

 

 

CAPILLARY EXCHANGE

Only the blood in capillaries is involved in exchanging material with tissue cells. Substances needed by cells move through the endothelial walls, into interstitial fluid, and then into tissue cells. Wastes move in the opposite direction. Two things normally do not leave the blood—large proteins and formed elements. Movement in and out of capillaries occurs by these methods:

   1. Diffusion—always from greater concentration to lesser concentration.

      a. Simple diffusion—this is the single most important method by which substances enter and leave capillaries. Simple diffusion can occur across endothelial cell plasma membranes, or through intercellular clefts and fenestrations.

      b. Facilitated diffusion—some substances also move through the endothelial plasma membrane this way.

 

 Substances which diffuse include:

                 Oxygen                         Amino acids

                 Carbon dioxide             Steroid hormones 

                 Glucose

In most capillaries, almost all plasma solutes diffuse EXCEPT large proteins. In sinusoids, intercellular clefts are very large so that even proteins & blood cells can enter the blood.

 

Remember the blood-brain barrier (BBB). Brain capillaries do not have pores or intercellular clefts. Their endothelial cells are connected by tight junctions. Special facilitated diffusion mechanisms move some nutrients and other substances into brain tissue. Some larger molecules can't enter at all.

 

   2. Transcytosis—this is a form of pinocytosis (vesicular transport). It is uncommon (not may things are moved this way). A few large nonlipid-soluble molecules are taken in by endocytosis on one side of the endothelial cells, pass through the cell, and leave on the other side by exocytosis.

              Insulin enters blood

              Antibodies crossing placenta

 

   3. Bulk flow—passive process in which large numbers of ions, molecules, or particles which are dissolved or suspended in fluid move together in the same direction. Pressure causes the movement, which is from an area of higher pressure to an area of lower pressure. The movement is faster than diffusion.

 

Bulk flow moves fluids and solutes both ways,  both in and out of capillaries.

      filtration—movement out of capillaries into tissue spaces

      reabsorption—movement from tissue spaces back into capillaries

 

Bulk flow:

   1. Helps deliver oxygen, nutrients, ions, etc. to cells

   2. Helps remove carbon dioxide, wastes, secretions, etc. from cells

   3. Helps regulate the relative volumes of blood and interstitial fluid (this is the most important function)

 

 

FORCES INVOLVED IN BULK FLOW

 

First, remember that wherever proteins are found, they have a strong tendency to draw fluid to themselves and hold it—this is called colloid osmotic pressure.

 

Forces encouraging filtration  (these forces predominate at the arterial end of a capillary):

   **1. Blood hydrostatic pressure (BHP) or just plain blood pressure

     2. Interstitial fluid osmotic pressure—this is normally a very small factor, since normal interstitial fluid contains very little protein

 

Forces encouraging reabsorption  (these forces predominate at the venous end of a capillary):

   **1. Blood colloid osmotic pressure—as fluid and other solutes leave, the remaining blood proteins become concentrated, giving a strong pull for fluid to return to the capillary.

     2. Interstitial fluid hydrostatic pressure—this would push fluid back into the capillary, but since this pressure is normally very low (often zero) this is a very  minor factor.

 

On the average, the volume of fluid reabsorbed should be almost the same as the

volume of fluid filtered. This is known as Starling’s Law of the Capillaries. In fact, reabsorption does not quite equal filtration, so the remaining 10 - 15% of the filtered fluid that is not reabsorbed into blood capillaries must be picked up and returned to the bloodstream by the lymphatic system (about 3liters/day).

 

EDEMA

If filtration greatly exceeds reabsorption, the result is an accumulation of fluid in tissue spaces, which is known as edema. Edema can be caused by:

v     Excess filtration:

·        Increased blood pressure causes greater volumes to be filtered

·        Increased permeability of capillaries allows excess filtration of fluid and also the escape of plasma proteins into tissue spaces

v     Inadequate reabsorption:

·        Decreased plasma protein levels due to malnutrition, burns, kidney disease, etc. lead to decreased blood colloid osmotic pressure

v     Blockage of normal lymphatic flow will also lead to edema

 

 

HEMODYNAMICS—FACTORS AFFECTING BLOOD FLOW TO TISSUES

 

 

VOLUME OF BLOOD FLOW (This does not refer to the total blood volume---it refers to the amount of blood moving through tissues.)

Cardiac output is the volume of blood that circulates through the systemic or pulmonary blood vessels each minute. Cardiac output equals stroke volume times

 

rate.  The 2 additional factors that influence how the cardiac output ecomes distriuted to the various tissues are Blood pressure and Resistance.

 

BLOOD PRESSURE—force exerted by blood on blood vessel walls. Blood pressure is generated by contraction of the ventricles and is greatest in the aorta—about 110/70 in a healthy young adult at rest. Mean arterial blood pressure is the average pressure and is calculated as follows:

          Diastolic BP  +  1/3 (Systolic BP  -  Diastolic BP)   =  MABP

                70           +                       1/3 (110 – 70)         =      83                    

If resistance does not change, increased cardiac output means increased MABP, and decreased cardiac output  causes decreased MABP. Blood in the cardiovascular system always flows from greater pressure to lesser pressure. Blood pressure falls steadily as blood flows through the aorta to large arteries, medium arteries,  small arteries, arterioles, capillaries, venules, and veins. The blood vessels with the lowest pressure are the venae cavae, and pressure is near  zero in the right atrium. There MUST be a pressure gradient for blood flow to continue.

 

RESISTANCE—opposition to blood flow, mainly due to friction between blood and the walls of blood vessels. Factors:

 

   1. Size of the lumen of blood vessels---this is the one that we adjust and change most readily. Other factors in resistance usually are fairly constant, or change gradually. Arterioles can change their size the most---minimum size is about 1/3 of the maximum. The smaller the lumen of a blood vessel, the greater the resistance to blood flow.

      a. Vasoconstriction---smooth muscle in the blood vessel wall contracts and the lumen becomes smaller. This increases resistance.

      b. Vasodilation---smooth muscle relaxes and lumen enlarges. This decreases resistance.

 

   2. Blood viscosity—“thickness” of blood. Depends mostly on number of RBC compared to the volume of plasma. To a smaller extent, it also depends on the concentration of plasma proteins. The greater the viscosity, the greater the resistance.

      a. Causes of increased viscosity: Dehydration (which decreases volume of plasma), production of unusually large numbers of RBC (polycythemia). If viscosity increases, blood pressure must increase to maintain flow.

      b. Causes of decreased viscosity: Decreased plasma protein levels, decreased RBC due to anemia or hemorrhage. This “thin” blood flows more easily, so blood pressure can drop and still deliver the same flow of blood to tissues.        

 

   3. Total blood vessel length—the greater the length, the greater the resistance, so the longer the total length of all the body’s blood vessels, the higher the blood pressure must be to push the blood through the all the vessels and back to the heart. Obesity causes increased resistance as the total length of the body’s blood vessels increases to supply the adipose tissue (almost 200 extra miles of blood vessels for each pound of fat).

 

  

SVR—Systemic vascular resistance  OR    TPR—Total peripheral resistance

Both of these terms refer to all the vascular resistance offered by all the systemic blood vessels. By vasodilation and vasoconstriction, arterioles (and other vessels to a lesser extent) change the SVR and therefore the blood pressure. The center for regulation of SVR is the vasomotor center of the medulla.

 

VENOUS RETURN

Blood flow in systemic veins is due to a pressure differential, with steadily lower pressures as the blood gets nearer the heart. The movement of skeletal muscles and respiratory muscles aids in venous return, by alternately squeezing and relaxing around the veins. Once blood has moved forward in a vein, valves prevent backflow. 

 

 

VELOCITY (SPEED) OF BLOOD FLOWmeasured in centimeters per second. Velocity in blood vessels is faster where the total cross-sectional area is smaller and slower where the total cross-sectional area is greater. The cross-sectional area in the above statement refers to the total cross-sectional area of ALL blood vessels of the type added together, not the size of any one single vessel. See chart (you do NOT have to memorize these values!)

 

TYPE OF VESSEL

CROSS-SECTIONAL AREA

(SQ  CM)

AVERAGE VELOCITY

(CM/SEC)

AORTA

3 - 5

40

CAPILLARIES

4500 - 6000

0.1

VENAE CAVAE

14

15

 

As seen above, velocity is fastest in the aorta and slowest in the capillaries, which allows the maximum time for capillary exchange to occur. The total time in a capillary is still not great—about 1-2 seconds on average. Speed picks up in veins as the blood flows back toward the heart, but never reaches maximum velocity.

 

Circulation time is the time required for a drop of blood to pass from the right atrium, through the pulmonary circulation, back to the left atrium, through the systemic circulation to the foot, and back to the right atrium. At rest, this should be about one minute.

 

 

 

CONTROL OF BLOOD PRESSURE AND BLOOD FLOW

 

Negative feedback systems regulating several functions work together to maintain blood pressure:

     Heart rate

     Stroke volume

     Systemic vascular resistance (diameter of blood vessels)

     Blood volume

 

Keep in mind:

   Increased blood volume, increased cardiac output & vasoconstriction cause INCREASED BP

   Decreased blood volume, decreased cardiac output & vasodilation cause DECREASED BP

 

1. Cardiovascular center--groups of neurons in the medulla oblongata regulate heart rate, force of contraction and blood vessel diameter. The CV center receives input from sensory receptors and higher brain regions. This includes input from:

      Baroreceptors which monitor pressure in certain blood vessels and the atria

      Chemoreceptors which monitor blood pH, CO2 level, and O2 level

      Proprioceptors which monitor movement of joints & muscles

      Cerebral cortex, limbic system, hypothalamus

After evaluating all this information, one or more of these areas within the CV center become active:

   a. Cardiostimulatory center—cardiac accelerator nerves carry sympathetic impulses to the heart, causing an increase in rate and force

   b. Cardioinhibitory center--sends parasympathetic impulses to the heart on the vagus nerve to decrease heart rate

   c. Vasomotor center--vasomotor nerves carry sympathetic impulses only (no parasympathetic) to smooth muscle in the walls of blood vessels. These affect all larger vessels but arterioles most of all. A moderate amount of vasoconstriction is maintained at all times (vasomotor tone). Increased sympathetic impulses cause vasoconstriction in most small arteries and arterioles, raising the BP. In skeletal muscle and the heart these same impulses cause vasodilation (remember fight-or-flight). In veins, sympathetic stimulation causes constriction that moves blood from the reservoirs into the circulation. Even though some vessels dilate, more constrict, so the net response is a rise in BP.

 

 

 

 

Blood vessels in:

Skeletal muscle

Heart

Lungs

Blood vessels in:

Skin

Viscera (such as digestive organs)

Overall effect

Blood pressure

Medium sympathetic stimulation

Medium diameter

Medium diameter

Vasomotor tone

Normal

Increased sympathetic stimulation (such as in physical activity)

Dilate

Constrict

Vasoconstriction

Rises

Decreased sympathetic stimulation (“Housekeeping”)

Constrict

Dilate

Vasodilation

Drops

 

 

2. Neural regulation

     a. Baroreceptor reflexes—baroreceptors are nerve cells that detect changes in pressure, located in the walls of arteries, veins and the right atrium. They act as receptors for these negative feedback systems:

         1) Carotid sinus reflex--purpose is to maintain sufficient blood flow to the brain. Baroreceptors located in the carotid sinuses, widenings of the proximal internal carotid arteries, detect a drop in BP to the brain and send impulses to the CV center of the medulla. These result in a quick increase in sympathetic impulses to raise BP by increasing both rate and force of heartbeat and by vasoconstriction. Although this reflex may be needed at various times, it is especially important when we suddenly sit or stand up from a reclining position. If the reflex does not operate smoothly, we have a brief feeling of "blacking out."

 

Baroreceptors can also report an increase in pressure. In this case, the cardiovascular center responds by causing a decrease in heart rate and force and vasodilation.

 

         2) Aortic reflex--baroreceptors in the aorta are concerned with general systemic blood pressure.

 

   b. Chemoreceptor reflexes—chemoreceptors are nerve cells that monitor the chemical content of the blood. They are located in the carotid bodies and the aortic bodies and detect changes in the blood levels of O2, CO2, and blood pH. If O2 or pH drop or if CO2 rises, these chemoreceptors send impulses that result in sympathetic stimulation to arterioles and veins. (Rate of breathing is also adjusted.)

 

3. Hormonal regulation

   a. Renin-angiotensin-aldosterone (RAA) system—when blood flow through the kidney decreases, special cells there secrete an enzyme, renin. This begins a series of steps that result in the production of the hormone angiotensin II, which raises BP in two ways:

      1) Causes vasoconstriction (fast)

      2) Stimulates secretion of aldosterone, which increases reabsorption of sodium and water in the kidney and therefore raises blood volume (slower)

   

   b. Epinephrine and norepinephrine (adrenal medulla) increase cardiac output and cause vasoconstriction of abdominal and cutaneous vessels; vasodilation of vessels of heart and skeletal muscle.

 

   c. Antidiuretic hormone (ADH)--causes vasoconstriction (also known as vasopressin). If ADH release is inhibited, vasodilation results.

 

   d. Atrial natriuretic peptide--produced by cells in the atria. It lowers BP by causing vasodilation and promoting loss of salt and water in the urine, lowering blood volume.

 

 

AUTOREGULATION OF BLOOD PRESSURE

 

In addition to changes in the systemic blood pressure, blood flow in a particular area of the body can also be regulated. This is known as autoregulation.

   a. Physical changes such as warming or cooling a particular body part--warming causes vasodilation, cooling causes vasoconstriction

   b. Chemical changes such as low oxygen levels, high CO2 levels, stretch of tissue and others can cause cells in the area to release chemicals called vasoactive factors. These influence diameter of blood vessels just in the affected area.

 

 

PULSE

The pulse is a pressure wave that travels through arteries. It is created by the alternate expansion and recoil of the elastic arteries after each left ventricular systole. It is best felt in arteries located near the surface of the body over a bone or other firm tissue.

   Radial artery at wrist

   Temporal artery lateral to orbit

   Common carotid artery lateral to larynx

Normal resting pulse 70 - 80 beats per minute.

Tachycardia—resting rate over 100

Bradycardia—rate under 60

 

BLOOD PRESSURE

 

Blood pressure is measured with a sphygmomanometer, usually in the brachial artery of the arm. Remember the definition, BP is the force exerted by the blood on the inner walls of the heart and blood vessels.

 

   Systolic BP (higher)--force following ventricular systole

   Diastolic BP (lower)--force during ventricular relaxation

 

In an average healthy young adult at rest, the BP might be 110/70.

 

Pulse pressure is the difference between systolic and diastolic pressures—for the 110/70 person it would be 40. Systolic/diastolic/pulse pressures should have approximately a 3 : 2 : 1 ratio.

 

SHOCK   

Shock is the failure of the cardiovascular system to deliver adequate amounts of oxygen and nutrients to the tissues (inadequate tissue perfusion). Cells try to go on, using anaerobic means of energy production. This results in accumulation of lactic acid and further damage.

TYPES OF SHOCK

 

1. Hypovolemic shock---decreased blood volume due to loss of blood                                      (hemorrhage) or fluid loss from vomiting/diarrhea, dehydration, etc.

 

2. Cardiogenic shock---inadequate heart function---myocardial infarction, heart failure, severe dysfuntion of valves

 

3. Vascular shock--widespread inappropriate vasodilation

    a. Anaphylactic shock—severe allergic reaction in which very large amounts of histamine are released

    b. Neurogenic shock—damage to cardiovascular center of the brain

    c. Septic shock—response to bacterial toxins

 

4. Obstructive shock---blockage of blood flow at any site in the CV system---pulmonary embolism, cardiac tamponade, etc.

 

 

STAGES OF SHOCK

 

I. COMPENSATED SHOCK  (NONPROGRESSIVE SHOCK)---minimal symptoms and the CV  system is able to compensate---loss of up to 10% total blood volume. Negative feedback systems restore homeostasis:

 

   1. Decreased blood flow to the kidneys causes activation of the renin-angiotensin-aldosterone system. Angiotensin II, a powerful natural vasopressor substance, causes vasoconstriction and the secretion of aldosterone

 

   2. Secretion of ADH, which also causes vasoconstriction and increases reabsorption of water by the kidneys.

 

   3. Baroreceptors report lowered BP and cause activation of sympathetic impulses which cause:

       a. Vasoconstriction of arterioles, esp. to skin and abdominal viscera

       b. Increased in heart rate and force of contraction

       c. Secretion of epinephrine/norepinephrine, which intensify the effects of a & b

 

   4. In response to hypoxia, affected cells produce local vasodilation and help restore oxygen to affected tissues. This may help a certain tissue, but overall it may interfere with efforts to raise systemic BP.

 

 

II. DECOMPENSATED SHOCK (PROGRESSIVE SHOCK )---shock becomes steadily worse as compensatory mechanisms cannot keep up---occurs with loss of 10-20% blood volume---positive feedback cycles arise:

 

   1. Depression of cardiac activity due to hypoxia of cardiac muscle---if systolic BP falls below 60, insufficient oxygen reaches the heart muscle---this causes the heart to pump with less force. This causes further hypoxia

 

   2. Depression of vasoconstriction occurs as the vasomotor center becomes depressed by lack of oxygen

 

   3. Increased permeability of capillaries causes a further drop in blood volume as fluid leaks out

 

   4. Intravascular (inappropriate) clotting due to sluggish circulation

 

   5. Cellular destruction, including the heart muscle cells

 

   6. Acidosis---dysfunctional cells release lactic acid

 

In this stage, medical intervention can reverse the changes.

 

 

III. IRREVERSIBLE SHOCK---rapid deterioration of the CV system that cannot be helped by compensatory mechanisms or medial intervention. ATP of the heart muscle is depleted and the heart cannot make more, so contractions cease.

 

 

SIGNS OF SHOCK

1. Hypotension—systolic BP below 90 due to vasodilation and

 decreased cardiac output

2. Clammy, cool, pale skin due to vasoconstriction of skin vessels

3. Sweating due to increased sympathetic stimulation

4. Reduced urine formation due to hypotension and increased aldosterone

   and ADH secretion

5. Cerebral ischemia causes altered mental status

6. Acidosis due to buildup of lactic acid

7. Tachycardia due to sympathetic stimulation and epinephrine release

8. Weak rapid pulse due to generalized vasodilation and reduced cardiac output

9. Thirst due to loss of extracellular fluid

10. Nausea due to impaired circulation to the digestive system

 

 

 

 

CIRCULATORY ROUTES

 

From birth onward, blood follows 2 basic routes:

 

   1. Systemic circulation--includes all blood vessels carrying blood from the left ventricle to body tissues except for lungs and back to the right atrium

   2. Pulmonary circulation--from right ventricle through the lungs and back to left atrium

 

 

SYSTEMIC CIRCULATION

 

All systemic arteries branch off the aorta:

   1. Ascending aorta--emerges from the left ventricle and passes upward behind the pulmonary trunk, gives off 2 coronary arteries that penetrate into heart muscle

   2. Arch of the aorta--gives off vessels that supply the head and arms

   3. Descending aorta--lies close to the backbone

      a. Thoracic aorta--travels through the thorax and penetrates diaphragm

      b. Abdominal aorta--continues through the abdominal cavity to the level of the 4th lumbar vertebra where it bifurcates to form the 2 common iliac arteries

 

Each section of the aorta gives off branches that supply the systemic tissues.